Wall (lasiommata megera) and Large Wall (lasiommata maera)

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2008 photos highlighted green. Click on any photo to go to an enlarged picture, or simply scroll down the page.

7900_male_Isere_5Jul07 - maera 8044_male_Isere_6Jul07 - maera 12324_male_Isere_9Jul08 - maera
13031_male_Valais, Switzerland_16Jul08 - maera 9279_female_Var_28Aug07 - maera 2225_female_Drome_1Jul06 - maera
10376_male_Alpes Maritimes_9May08 - megera 6678_male_Alpes Maritimes_8Jun07 - megera 7144_female_Var_18Jun07 - megera
   
7077_female_Var_16Jun07 - megera    

Megera is declining alarmingly in the UK but still reasonably common in southern France, although maera sometimes seems commoner, especially at medium and high altitudes. Megera emerges very early in the season and again in October when it seems to be quite common, in places very common, in Var. Maera often poses quite obligingly for photographs, although it has a habit of settling in the dark shady areas on walls and rocks, presumably to escape the heat in the middle of the day. The nominate form of maera is easy to differentiate from megera as it is almost uniformly dark brown (not to mention maera usually being appreciably larger). However, the female of the southern form adrasta, which is said (in H&R) to be the predominant form in southern France and Iberia, has lighter markings and is more extensively orange, making differentiation from megera more difficult. I have seen adrasta in the Pyrenees, but on balance I believe the maera in Var (as in 9279) are of the nominate form, so I wonder if H&R is right on this point.

 

The illustrations of megera in T&L show the female to be very similar to the male, with heavier dark brown bands breaking up the orange as in 10376. This originally led me to believe that 7077 and 7144 were maera of the form adrasta as they were much closer to the T&L illustrations of adrasta than to the female megera.

On checking H&R and the Lafranchis ID book, the illustrations of female megera there are much closer to 7077 and 7144, so I am left to consider that the T&L illustration is potentially quite misleading. My thanks to Tim Cowles whose detailed analysis points up the key differentiators, including the underside where three of the maera ocelli (s2, s3 and s6) are appreciably larger (clear in 2225 and to a lesser extent in 13031), whereas the six megera unh ocelli are of broadly equal size (as in 6678). You don't tend to find invaluable information like this in books. For simplicity and comparison, I decided to put megera and maera on the same page.

 

There is also the Northern Wall Brown (L. petropolitana) which is very similar to the nominate form of maera, which belies its name as it is found in Scandinavia and the Alps and Pyrenees, but not in between. Petropolitana is slightly smaller and darker and has a clearer, although sometimes not very distinctive, discal line on the uph.

 

7900: a typical male maera of the nominate form. Altitude 1450m.

8044: a male maera of the nominate form. Altitude 1500m.

12324: a male maera underside of the nominate form.

13031: a male maera underside of the nominate form.

9279: a female maera of the nominate form, not quite orange enough to suggest the form adrasta. Altitude 1500m.

2225: female(?) maera: there is a large main unf apical ocellus, plus a second, smaller, fused ocellus, and a mini-ocellus attached above, a full set of unh post-discal rings (T&L shows the male with a reduced ocellus in s4 - is this indicative?), light unh ground colour and relatively weak discal lines, all indicating female. I suspect it is the nominate form, not adrasta. Altitude 1180m.

10376: a typical male megera. A difficult butterfly to approach and photograph, I find.

6678: a male megera: the underside carries several suggestions of maera, notably the number of rings in each ocellus, the general light colouring, and the large unf ocellus, but the consistency of the size of the unh ocelli is a sure pointer to megera, as noted above. Altitude 330m.

7144: a female megera, quite extensively marked orange, which, in many respects, looks remarkably like the T&L illustration of the female maera of the southern form adrasta. However, the orange in the uph post-discal region is a pointer toward megera. Altitude 780m.

7077: a female megera. It is exceptionally orange and looks nothing like the illustration in T&L. Altitude 220m.

 

7900_male_Isere_5Jul07 - maera

 

8044_male_Isere_6Jul07 - maera

 

12324_male_Isere_9Jul08 - maera

 

13031_male_Valais, Switzerland_16Jul08 - maera

 

9279_female_Var_28Aug07 - maera

 

2225_female_Drome_1Jul06 - maera

 

10376_male_Alpes Maritimes_9May08 - megera

 

6678_male_Alpes Maritimes_8Jun07 - megera

 

7144_female_Var_18Jun07 - megera

 

7077_female_Var_16Jun07 - megera